In 2004, I upgraded my G4/450 Sawtooth with a PowerLogix 1.2Ghz (single) G4 chip with 2MB level 3 cache, and it was a great improvement and very stable. I was running only OS9.2.2 at the time.
The only reason I mention that here, is that once I forgot, and zapped the PRAM. PowerLogix specifically warned against zapping the PRAM after installing the upgrade CPU card, and lo and behold, I had to REINSTALL the OLD CPU card, THEN zap the PRAM again, and DOWNGRADE the Firmware, with the included PowerLogix CD-ROM. THEN put the new CPU card back in, and then re-upgrade the Firmware, and all was fine again.
Eventually, I installed Tiger and all was well, but last week, I was trying to reinstall a clean OS9 before I gave the computer to a friend, and when OS9 wouldn't install, I somehow got 1/2 way through the Firmware downgrade, and the keyboard wasn't recognized, and I had to abort. The computer has never booted since then.
My point is that it is possible to corrupt the Boot ROM during Firmware upgrading or downgrading, which may make booting impossible, or temporarily so; and this impacts differently on OS9 or OSX.
Welcome to The Twilight Zone.
